A Functional Biology of Free-Living Protozoa by Johanna Laybourn-Parry BSc, MSc, PhD (auth.)

By Johanna Laybourn-Parry BSc, MSc, PhD (auth.)

General Editor: Peter Calow, division of Zoology, college of Sheffield, England the most goal of this sequence can be to demonstrate and to give an explanation for the way in which organisms 'make a residing' in nature. on the middle of this - their func­ tional biology - is the best way organisms collect after which utilize assets in metabolism, stream, progress, copy, etc. those procedures will shape the elemental framework of the entire books within the sequence. every one e-book will be aware of a selected taxon (species, family members, type or maybe phylum) and may compile info at the shape, body structure, ecology and evolutionary biology of the crowd. the purpose could be not just to explain how organisms paintings, but additionally to contemplate why they've got come to paintings in that means. by way of focusing on taxa that are renowned, it really is was hoping that the sequence won't basically illustrate the luck of choice, but additionally convey the restrictions imposed upon it via the physiological, morphological and developmental limita­ tions of the teams. one other very important characteristic of the sequence can be its organismic orienta­ tion. every one e-book will emphasise the significance of useful integra­ tion within the day by day lives and the evolution of organisms. this is often the most important in view that, although it can be actual that organisms may be regarded as collections of gene-determined features, they however have interaction with their atmosphere as built-in wholes and it really is during this context that specific qualities were subjected to traditional choice and feature evolved.

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13). Frequently other individuals of Actinophrys may fuse with the feeding individual, separating again on completion of digestion. This represents a useful adaptation allowing cooperation in the feeding on, and processing of, a prey too large for a single heliozoan to cope with. 13: A Heliozoan Ingesting a Ciliate Prey. Source: Based on Patterson and Hausmann (1981). Foraminiferans also exploit a wide range of food. Some, for example Mansipella arenaria, are suspension feeders taking small particles including bacteria, algae and fine detritus.

The rodorgan is composed of a pair of rods, each constructed from 100-200 microtubules, enclosed in a sheathing membrane anteriorly (Nisbet, 1974). Peranema feeds on a variety of food organisms, including bacteria and detritus, but can also successfully ingest food items as large as itself. When feeding on Euglena, for example, the rodorgan and part of the adjacent region of the ventral body surface move forwards until it touches the prey. The rodorgan protrudes and becomes attached to Euglena, while the body 44 How Protozoa Obtain Energy of Peranema then moves forwards and the prey moves into the expanded cytostome.

The Golgi apparatus, if present, may participate in the condensation of enzymes (Muller, 1967). Hydrolytic enzymes appear to fall into two functional groups in Protozoa, those which are engaged in digestive processes occurring within the cell and others which function in the vacuoles. Muller (1967) has speculated on the evolution of digestive processes in Protozoa. The widely-accepted view is that intracellular digestion is the ancient form of food breakdown and the extracellular cavity digestion of food as seen in metozoans evolved from intracellular digestion only after the formation of multicellular organisms.

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